The Celestial Lions Space Marines are being targeted by the Inquisition and have been brought to the brink of extinction. Will they regroup and rebuild, or will they choose to go out in a final blaze of glory? That is what Grimaldus must decide...

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As if the prospect of Black Templars and Celestial Lions fighting side-by-side against an ork horde wasn't enough, Blood And Fire also shines a light on some of the deep divisions within the Imperium. in the face of such inner turmoil and underhanded treachery, even the zeal of the legendary Grimaldus may not be enough to save his Space Marine allies.